To reproduce:
1. Open attached project;
2. Open "Main" scene;
3. Enter Play mode;
-Notice particle system works.
4. Exit Play mode;
5. Build and Run the project.

Expected result: Particle System works in both Editor and Build.
Actual result: Particle System does not work on Build.

Note: If VR is disabled, Particle System works on Build too.

Reproduced on: 5.6.1p4, 5.6.2f1, 2017.1.0f2, 2017.2.0b1.
Not reproduced on: 5.6.1f1, 5.6.1p2, 2017.1.0f1.
Regression introduced on: 5.6.1p3, 2017.1.0f2.

FIXED 2018.3.1f1

Note: on versions 2017.2.0b1, 2017.2.0b2 almost impossible to use Editor, when entered Play mode.