Animations get skewed/distorted when animated objects are scaled and "Optimize Game Objects" is selected

How to reproduce:

1. Open the attached project
2. Open and play "0-4 Weak Floors and Switch" scene
3. Notice that animation is distorted

Workaround:

1. Do not scale the animated object (any scaling would be done in the model)
2. Disable "Optimize Game Objects"
3. Select all distorted animations and change its Animation Compression from Keyframe Reduction to Off

Reproducible: 5.3.0f4, 5.3.1p1, 5.3.1p3
Not reproducible: 5.2.4f1