How to reproduce:
1. Open the attached “IN-95200_align_to_direction“ project
2. Open the “SampleScene“ and enter Scene View
3. Select “Particle System“ GameObject
4. In the Inspector window, under the Particle System component, go to the “Shape“ module, set the Scale of the Z-axis to 0 and ensure that “Align To Direction“ is enabled
5. Observe the result

Expected result: Particles are rendered
Actual result: Particles are not rendered

Reproducible with: 2022.3.58f1, 6000.0.38f1, 6000.1.0b6, 6000.2.0a3

Reproducible on: MacOS 15.3 (M4), Windows 11 (by user)
Not reproducible on: No other environments tested

Notes:
- Reproducible in Player
- All available shapes under the Shape module were tested and the issue was reproduced only with the Box shape and Rectangle shape