- How to reproduce in the editor -

1. Open "Main" scene.
2. In the project window, select asset "Assets/AssetBundleResources/SimpleMesh.fbx"
3. Make sure "Read/Write Enabled" parameter to false.
4. Build Asset Bundles (menu Assets/AssetBundles/Build AssetBundles)
5. Launch local server (menu Assets/AssetBundles/Local AssetBundle Server)
6. Enter play mode
- notice how the particle system in the instantiated prefab doesn't show any particle.
7. Quit play mode.
8. In the project window, select asset "Assets/AssetBundleResources/SimpleMesh.fbx"
9. Change "Read/Write Enabled" parameter to true.
10. Build Asset Bundles again.
11. Enter play mode.
- notice how the particle system now works properly.

- How to reproduce in a build -

1. Open "Main" scene (in the editor).
2. In the project window, select asset "Assets/AssetBundleResources/SimpleMesh.fbx"
3. Make sure "Read/Write Enabled" parameter to false.
4. Build Asset Bundles (menu Assets/AssetBundles/Build AssetBundles)
5. Launch local server (menu Assets/AssetBundles/Local AssetBundle Server)
6. Build & Run
- notice how the build crashes after the unity logo animation (present only in 5.3.5p2, in 5.4 did not crash, but result was same as in editor)

7. Back in the editor, project window, select asset "Assets/AssetBundleResources/SimpleMesh.fbx"
8. Change "Read/Write Enabled" parameter to true.
9. Build Asset Bundles again.
10. Build & Run
- it should work fine.

Expected result: It should be possible for particle system to use mesh which is in another asset bundle
Actual result: Particle system is invisible if mesh is loaded from another bundle

Reproduced on:
Crashes on 5.1.4f1, 5.2.4f1, 5.3.5p2, On 5.4.0b21 it does not crash, but particles are still invisible